TN2009000512A1 - Modifying glycoprotein production in plants - Google Patents

Modifying glycoprotein production in plants

Info

Publication number: TN2009000512A1
Authority: TN; Tunisia
Prior art keywords: plant; nucleotide sequence; protein; interest; plants
Prior art date: 2007-06-15

Application number

TNP2009000512A

Other languages

English (en)

Inventor

Aoust Marc-Andre D

Estelle Marquet-Blouin

Muriel Bardor

Carole Burel

Loic Faye

Patrice Lerouge

Louis-Philippe Vezina

Véronique Gomord

Stéphanie Aquin

Christophe Rihouey

Thomas Paccalet

Christophe Sourrouille

Original Assignee

Centre Nat Rech Scient

Medicago Inc

Univ Rouen

Priority date (The priority date is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the date listed.)

2007-06-15

Filing date

2009-12-10

Publication date

2011-03-31

2009-12-10 Application filed by Centre Nat Rech Scient, Medicago Inc, Univ Rouen filed Critical Centre Nat Rech Scient

2011-03-31 Publication of TN2009000512A1 publication Critical patent/TN2009000512A1/fr

Classifications

- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8241—Phenotypically and genetically modified plants via recombinant DNA technology
- C12N15/8242—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits
- C12N15/8257—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits for the production of primary gene products, e.g. pharmaceutical products, interferon
- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8216—Methods for controlling, regulating or enhancing expression of transgenes in plant cells
- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8216—Methods for controlling, regulating or enhancing expression of transgenes in plant cells
- C12N15/8218—Antisense, co-suppression, viral induced gene silencing [VIGS], post-transcriptional induced gene silencing [PTGS]
- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8241—Phenotypically and genetically modified plants via recombinant DNA technology
- C12N15/8242—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits
- C12N15/8243—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits involving biosynthetic or metabolic pathways, i.e. metabolic engineering, e.g. nicotine, caffeine
- C12N15/8245—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits involving biosynthetic or metabolic pathways, i.e. metabolic engineering, e.g. nicotine, caffeine involving modified carbohydrate or sugar alcohol metabolism, e.g. starch biosynthesis
- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8241—Phenotypically and genetically modified plants via recombinant DNA technology
- C12N15/8242—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits
- C12N15/8257—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits for the production of primary gene products, e.g. pharmaceutical products, interferon
- C12N15/8258—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits for the production of primary gene products, e.g. pharmaceutical products, interferon for the production of oral vaccines (antigens) or immunoglobulins
- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
- C12N9/10—Transferases (2.)
- C12N9/1048—Glycosyltransferases (2.4)
- C12N9/1051—Hexosyltransferases (2.4.1)

Landscapes

Health & Medical Sciences (AREA)
Genetics & Genomics (AREA)
Life Sciences & Earth Sciences (AREA)
Engineering & Computer Science (AREA)
Chemical & Material Sciences (AREA)
Molecular Biology (AREA)
Biotechnology (AREA)
Organic Chemistry (AREA)
Zoology (AREA)
Wood Science & Technology (AREA)
Bioinformatics & Cheminformatics (AREA)
Biomedical Technology (AREA)
General Engineering & Computer Science (AREA)
General Health & Medical Sciences (AREA)
Biochemistry (AREA)
Microbiology (AREA)
Cell Biology (AREA)
Plant Pathology (AREA)
Biophysics (AREA)
Physics & Mathematics (AREA)
Medicinal Chemistry (AREA)
Pharmacology & Pharmacy (AREA)
Nutrition Science (AREA)
Immunology (AREA)
Virology (AREA)
Preparation Of Compounds By Using Micro-Organisms (AREA)
Micro-Organisms Or Cultivation Processes Thereof (AREA)
Peptides Or Proteins (AREA)
Enzymes And Modification Thereof (AREA)
Breeding Of Plants And Reproduction By Means Of Culturing (AREA)
Cultivation Of Plants (AREA)

TNP2009000512A 2007-06-15 2009-12-10 Modifying glycoprotein production in plants TN2009000512A1 (en)

Applications Claiming Priority (2)

Application Number	Priority Date	Filing Date	Title
US94434407P	2007-06-15	2007-06-15
PCT/CA2008/001139 WO2008151440A1 (fr)	2007-06-15	2008-06-13	Modification de la production de glycoprotéine dans des plantes

Publications (1)

Publication Number	Publication Date
TN2009000512A1 true TN2009000512A1 (en)	2011-03-31

Family

ID=40129187

Family Applications (1)

Application Number	Title	Priority Date	Filing Date
TNP2009000512A TN2009000512A1 (en)	2007-06-15	2009-12-10	Modifying glycoprotein production in plants

Country Status (24)

Country	Link
US (1)	US20110008837A1 (fr)
EP (1)	EP2155880B1 (fr)
JP (1)	JP5783720B2 (fr)
KR (2)	KR101385050B1 (fr)
CN (3)	CN104962579A (fr)
AU (1)	AU2008261527C1 (fr)
BR (1)	BRPI0812538B1 (fr)
CA (2)	CA2795379A1 (fr)
DK (1)	DK2155880T3 (fr)
EG (1)	EG26600A (fr)
ES (1)	ES2624776T3 (fr)
HU (1)	HUE031698T2 (fr)
IL (1)	IL202568A (fr)
MA (1)	MA31516B1 (fr)
MX (1)	MX2009013665A (fr)
NZ (2)	NZ598417A (fr)
PL (1)	PL2155880T3 (fr)
PT (1)	PT2155880T (fr)
RU (2)	RU2499053C2 (fr)
SG (1)	SG185249A1 (fr)
SI (1)	SI2155880T1 (fr)
TN (1)	TN2009000512A1 (fr)
WO (1)	WO2008151440A1 (fr)
ZA (1)	ZA201000207B (fr)

Families Citing this family (18)

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AU2009202819B2 (en)	2007-07-13	2014-08-28	Aramis Biotechnologies Inc.	Recombinant influenza virus-like particles (VLPs) produced in transgenic plants expressing hemagglutinin
CA2615372A1 (fr)	2007-07-13	2009-01-13	Marc-Andre D'aoust	Particules semblables au virus grippal a comprenant de l'hemagglutinine
JP5809054B2 (ja)	2008-07-08	2015-11-10	メディカゴインコーポレイテッド	可溶性組換えインフルエンザ抗原
EP2318530B1 (fr)	2008-07-18	2016-06-22	Medicago Inc.	Nouvel épitope d immunisation contre l'influenzavirus
HUE039100T2 (hu)	2009-06-24	2018-12-28	Medicago Inc	Hemagglutinint tartalmazó kimer influenzavírus-jellegû részecskék
MY158475A (en)	2009-09-22	2016-10-14	Medicago Inc	Method of preparing plant-derived proteins
TWI620816B (zh)	2011-03-23	2018-04-11	苜蓿股份有限公司	植物衍生蛋白回收方法
US11673926B2 (en) *	2011-06-07	2023-06-13	Ibio, Inc.	In vivo de-glycosylation of recombinant proteins by co-expression with PNGase F
US9181532B2 (en)	2011-07-29	2015-11-10	Icon Genetics Gmbh	Production of galactosylated N-glycans in plants
FI2760882T3 (fi)	2011-09-30	2023-07-17	Medicago Inc	Viruksen kaltaisen partikkelin saantojen lisääminen kasveissa
WO2013169009A1 (fr) *	2012-05-08	2013-11-14	경상대학교 산학협력단	Plante pour la production de n-glycane riche en mannose, et procédé de production de n-glycane riche en mannose à l'aide de celle-ci
IL295155A (en)	2013-03-28	2022-09-01	Medicago Inc	Production of influenza virus-like particles in plants
US12467058B2 (en)	2013-03-28	2025-11-11	Aramis Biotechnologies Inc.	Influenza virus-like particle production in plants
ES2787042T3 (es)	2014-07-11	2020-10-14	Medicago Inc	Modificación de la producción de proteínas en plantas
CN107810271B (zh) *	2014-11-20	2021-09-28	耶路撒冷希伯来大学伊萨姆研发公司	用于在植物细胞中生产具有改变的糖基化模式的多肽的组合物和方法
WO2018098572A1 (fr)	2016-12-01	2018-06-07	Plantform Corporation	Plante transgénique à activité fucosyltransférase et xylosyltransférase réduite
CN111328347A (zh) *	2017-09-11	2020-06-23	R.J.雷诺兹烟草公司	通过与p21共表达来增加植物中感兴趣基因的表达的方法和组合物
JP7566045B2 (ja) *	2021-01-15	2024-10-11	デンカ株式会社	栽培条件を決定する方法、及び目的タンパク質又は目的ペプチドの製造方法

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GB9524350D0 (en) *	1995-11-29	1996-01-31	Lynxvale Ltd	Enhancer-increased gene expression in plants
US5945326A (en) *	1997-03-20	1999-08-31	New England Biolabs, Inc.	Method for cloning and producing the Spel restriction endonuclease
JP2002543760A (ja) *	1998-12-09	2002-12-24	達治関	ヒト型糖鎖をもつ糖タンパク質の生産方法
US7125978B1 (en) *	1999-10-04	2006-10-24	Medicago Inc.	Promoter for regulating expression of foreign genes
WO2001025455A2 (fr) *	1999-10-04	2001-04-12	Medicago Inc.	Promoteur destine a reguler l'expression de genes etrangers
AU1573601A (en) *	1999-10-21	2001-04-30	Monsanto Company	Post-translational modification of recombinant proteins produced in plants
ES2349427T3 (es) *	1999-10-26	2011-01-03	Stichting Dienst Landbouwkundig Onderzoek	Glicosilación de tipo mamífero en plantas.
CA2925231A1 (fr) *	1999-10-26	2001-05-03	Stichting Dienst Landbouwkundig Onderzoek	Glycosylation de type mammifere dans des plantes transgeniques par l'expression de galactosyltrasferases et de sialyl transferases
JP2005505234A (ja) *	2001-01-19	2005-02-24	ザ　ダウ　ケミカル　カンパニー	植物細胞によるヒト型糖鎖をもつ糖タンパク質の分泌生産方法
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PL224787B1 (pl)	2003-01-22	2017-01-31	Roche Glycart Ag	Wyizolowany kwas nukleinowy, ssaczy wektor ekspresyjny, fuzje polipeptydowe, komórki gospodarza, sposób wytwarzania fuzji polipeptydowych, sposoby in vitro i ex vivo modyfikowania profilu glikozylacji, wyizolowana cząsteczka kwasu nukleinowego
CN1761746B (zh) *	2003-01-22	2016-03-23	罗氏格黎卡特股份公司	融合构建体及其用来生产Fc受体结合亲和性和效应子功能提高的抗体的用途
BRPI0410340A (pt) *	2003-05-05	2006-05-30	Dow Agrosciences Llc	vetores e células para preparar composições imunoprotetoras derivadas de plantas transgênicas
JP2006212019A (ja) *	2004-04-30	2006-08-17	National Institute Of Agrobiological Sciences	植物を用いたユビキノン−１０の製造方法

2008
- 2008-06-13 PL PL08772802T patent/PL2155880T3/pl unknown
- 2008-06-13 CA CA2795379A patent/CA2795379A1/fr not_active Abandoned
- 2008-06-13 JP JP2010511461A patent/JP5783720B2/ja not_active Expired - Fee Related
- 2008-06-13 MX MX2009013665A patent/MX2009013665A/es active IP Right Grant
- 2008-06-13 ES ES08772802.8T patent/ES2624776T3/es active Active
- 2008-06-13 US US12/664,567 patent/US20110008837A1/en not_active Abandoned
- 2008-06-13 KR KR1020107000811A patent/KR101385050B1/ko not_active Expired - Fee Related
- 2008-06-13 KR KR1020137014893A patent/KR20130075787A/ko not_active Ceased
- 2008-06-13 WO PCT/CA2008/001139 patent/WO2008151440A1/fr not_active Ceased
- 2008-06-13 CN CN201510364867.XA patent/CN104962579A/zh active Pending
- 2008-06-13 BR BRPI0812538-4A patent/BRPI0812538B1/pt not_active IP Right Cessation
- 2008-06-13 SG SG2012066486A patent/SG185249A1/en unknown
- 2008-06-13 AU AU2008261527A patent/AU2008261527C1/en not_active Ceased
- 2008-06-13 CN CN2008801031743A patent/CN101918559A/zh active Pending
- 2008-06-13 SI SI200831699A patent/SI2155880T1/sl unknown
- 2008-06-13 NZ NZ598417A patent/NZ598417A/xx not_active IP Right Cessation
- 2008-06-13 NZ NZ581944A patent/NZ581944A/en unknown
- 2008-06-13 HU HUE08772802A patent/HUE031698T2/en unknown
- 2008-06-13 CA CA2700180A patent/CA2700180C/fr active Active
- 2008-06-13 EP EP08772802.8A patent/EP2155880B1/fr active Active
- 2008-06-13 CN CN2013101091944A patent/CN103382487A/zh active Pending
- 2008-06-13 PT PT87728028T patent/PT2155880T/pt unknown
- 2008-06-13 RU RU2010101024/10A patent/RU2499053C2/ru active
- 2008-06-13 DK DK08772802.8T patent/DK2155880T3/en active
2009
- 2009-12-07 IL IL202568A patent/IL202568A/en active IP Right Grant
- 2009-12-10 TN TNP2009000512A patent/TN2009000512A1/fr unknown
- 2009-12-14 EG EG2009121826A patent/EG26600A/en active
2010
- 2010-01-12 ZA ZA2010/00207A patent/ZA201000207B/en unknown
- 2010-01-12 MA MA32504A patent/MA31516B1/fr unknown
2013
- 2013-08-08 RU RU2013137208/10A patent/RU2013137208A/ru not_active Application Discontinuation

Also Published As

Publication number	Publication date
PT2155880T (pt)	2016-11-18
JP2010531136A (ja)	2010-09-24
ZA201000207B (en)	2013-09-25
CN104962579A (zh)	2015-10-07
AU2008261527B2 (en)	2014-04-10
NZ598417A (en)	2013-06-28
MX2009013665A (es)	2010-06-02
AU2008261527B8 (en)	2014-04-24
WO2008151440A1 (fr)	2008-12-18
CN103382487A (zh)	2013-11-06
EP2155880B1 (fr)	2016-08-10
MA31516B1 (fr)	2010-07-01
EG26600A (en)	2014-03-23
SG185249A1 (en)	2012-11-29
KR101385050B1 (ko)	2014-05-13
IL202568A0 (en)	2011-08-01
NZ581944A (en)	2012-03-30
PL2155880T3 (pl)	2017-06-30
CN101918559A (zh)	2010-12-15
AU2008261527A8 (en)	2010-01-21
CA2700180C (fr)	2013-01-29
AU2008261527A1 (en)	2008-12-18
SI2155880T1 (sl)	2017-03-31
EP2155880A4 (fr)	2010-08-04
CA2700180A1 (fr)	2008-12-18
RU2010101024A (ru)	2011-07-20
BRPI0812538A2 (pt)	2014-09-30
US20110008837A1 (en)	2011-01-13
CA2795379A1 (fr)	2008-12-18
BRPI0812538B1 (pt)	2021-03-16
RU2499053C2 (ru)	2013-11-20
KR20130075787A (ko)	2013-07-05
ES2624776T3 (es)	2017-07-17
JP5783720B2 (ja)	2015-09-24
HUE031698T2 (en)	2017-07-28
EP2155880A1 (fr)	2010-02-24
WO2008151440A8 (fr)	2010-03-04
KR20100037602A (ko)	2010-04-09
DK2155880T3 (en)	2016-12-05
AU2008261527C1 (en)	2014-09-04
IL202568A (en)	2015-02-26
RU2013137208A (ru)	2015-02-20
WO2008151440A4 (fr)	2009-02-26

Publication	Publication Date	Title
TN2009000512A1 (en)	2011-03-31	Modifying glycoprotein production in plants
PH12012500816A1 (en)	2017-07-26	Modified oil encapsulating proteins and uses thereof
SG10201803982TA (en)	2018-07-30	Antibodies and processes for preparing the same
MY160680A (en)	2017-03-15	Process for producing molecules containing specialized glycan structures
DE60022313D1 (de)	2005-10-06	Alternansucrase-kodierende nukleinsäure moleküle
DE50014279D1 (de)	2007-06-06	Nukleinsäuremoleküle aus pflanzen codierend enzyme, die an der stärkesynthese beteiligt sind
AR048024A1 (es)	2006-03-22	Plantas con actividad aumentada de distintas enzimas fosforilantes del almidon
MX2009009371A (es)	2009-09-21	Produccion mejorada de derivados de acidos grasos.
MX2009011751A (es)	2010-02-22	Produccion de proteinas de alto contenido de manosa en cultivo de plantas.
WO2008132314A3 (fr)	2008-12-31	Procede de synthese de nanoparticules metalliques cubiques en presence de deux reducteurs
DK1725666T3 (da)	2012-05-07	Planter med reduceret aktivitet for stivelsesphosphoryleringsenzymet phosphoglucan-vand-dikinase
WO2011100477A3 (fr)	2012-04-12	Anticorps et procédés de préparation de ceux-ci
CY1109471T1 (el)	2014-08-13	Μεθοδοι και μεσα για την παραγωγη υαλουρονανης
WO2009080743A3 (fr)	2009-12-30	Plantes à rendement accru (ko nue)
WO2010066411A3 (fr)	2010-09-16	Polypeptides à activité de cellobiohydrolase ii
MX2009010867A (es)	2009-10-28	Plantas con caracteristicas de crecimientos mejoradas en condiciones de disponibilidad reducida de nutrientes y un metodo para producirlas.
IN2012DN01454A (fr)	2015-06-05
EP2162537A4 (fr)	2010-07-14	Utilisation d'organismes genetiquement modifies pour generer des enzymes de decomposition de biomasse
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