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USPP10778P - `Cavalier` zoysiagrass plant - Google Patents

`Cavalier` zoysiagrass plant Download PDF

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Publication number
USPP10778P
USPP10778P US08/890,959 US89095997V US10778P US PP10778 P USPP10778 P US PP10778P US 89095997 V US89095997 V US 89095997V US 10778 P US10778 P US 10778P
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cavalier
zoysiagrass
sup
mean
meyer
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Milton Charles Engelke
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Texas A&M University
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Priority to PCT/US1998/014283 priority patent/WO1999002029A1/en
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    • A—HUMAN NECESSITIES
    • A01—AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01H—NEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
    • A01H6/00—Angiosperms, i.e. flowering plants, characterised by their botanic taxonomy
    • A01H6/46—Gramineae or Poaceae, e.g. ryegrass, rice, wheat or maize
    • A01H6/469—Zoysia
    • Y—GENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y10—TECHNICAL SUBJECTS COVERED BY FORMER USPC
    • Y10T—TECHNICAL SUBJECTS COVERED BY FORMER US CLASSIFICATION
    • Y10T83/00—Cutting
    • Y10T83/647—With means to convey work relative to tool station
    • Y10T83/6492—Plural passes of diminishing work piece through tool station
    • Y10T83/6499—Work rectilinearly reciprocated through tool station
    • Y10T83/65—With means to cause or permit angular re-orientation of work about axis parallel to plane of cut
    • Y—GENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y10—TECHNICAL SUBJECTS COVERED BY FORMER USPC
    • Y10T—TECHNICAL SUBJECTS COVERED BY FORMER US CLASSIFICATION
    • Y10T83/00—Cutting
    • Y10T83/647—With means to convey work relative to tool station
    • Y10T83/6492—Plural passes of diminishing work piece through tool station
    • Y10T83/6499—Work rectilinearly reciprocated through tool station
    • Y10T83/6508—With means to cause movement of work transversely toward plane of cut
    • Y10T83/6515—By means to define increment of movement toward plane of cut
    • Y10T83/6518—By pusher mechanism
    • Y10T83/6523—Including plural, simultaneously acting pusher elements
    • Y10T83/6526—With additional means to retract elements
    • Y—GENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y10—TECHNICAL SUBJECTS COVERED BY FORMER USPC
    • Y10T—TECHNICAL SUBJECTS COVERED BY FORMER US CLASSIFICATION
    • Y10T83/00—Cutting
    • Y10T83/929—Tool or tool with support
    • Y10T83/9372—Rotatable type
    • Y10T83/9403—Disc type

Definitions

  • the present invention relates to a new and distinct asexually reproduced variety of perennial zoysiagrass (Zoysia matrella (L.)) Merr.
  • This invention relates to a new and distinct perennial zoysiagrass cultivar identified as ⁇ Cavalier ⁇ zoysiagrass (hereinafter referred to as ⁇ Cavalier ⁇ ), that was tested as DALZ8507.
  • ⁇ Cavalier ⁇ was discovered in a cultivated area that received mowing and traffic on Seoul National University campus, Suwon, South Korea. It was identified as a unique turfgrass clone growing among a diverse population of common zoysiagrasses native to the area.
  • FIG. 1 depicts an up-close view of the leaf blade and ligule of ⁇ Cavalier ⁇ .
  • FIG. 2 depicts entire leaf blades and ligule of ⁇ Cavalier ⁇ .
  • FIG. 3 is a DNA fingerprint of ⁇ Cavalier ⁇ in comparison with known varieties of zoysiagrass.
  • ⁇ Cavalier ⁇ was characterized in greenhouse and field conditions.
  • ⁇ Cavalier ⁇ is a unique variety of zoysiagrass (Zoysia matrella (L.) Merr that was discovered under cultivated conditions described above.
  • ⁇ Cavalier ⁇ was asexually reproduced by cutting of stolons and rhizomes, rooting them in soil, and planting of the rooted material to provide planting stock for studying performance and for comparison of morphological characters after propagation.
  • ⁇ Cavalier ⁇ has been propagated by sod, plugs, sprigs, and stolons. Seed reproduction with self-fertility is not common in the Zoysia sp. No seedling establishment from ⁇ Cavalier ⁇ has been noticed in either greenhouse or field studies.
  • ⁇ Cavalier ⁇ spreads primarily by stolons. It has an intermediate to rapid growth rate, and has intermediate to low water use requirements. ⁇ Cavalier ⁇ produces little thatch with an optimum mowing height of 3/8" to 2.5". ⁇ Cavalier ⁇ will generally cover in 10-12 months growing time when plug planted with 3" ⁇ 4" plugs planted on 12" centers or by sprigs.
  • ⁇ Cavalier ⁇ has both rhizome and stolon growth.
  • the stolons of ⁇ Cavalier ⁇ have a mean internode length of 16.7 mm between the fourth to the fifth node, 18.4 mm between the second and third nodes, with a mean stolon width and diameter from 1.35 to 1.14 mm (Tables 1,2).
  • the stolons of ⁇ Cavalier ⁇ root adventitiously at the nodes. Color notations of plant tissues were based on the Munsell Color Charts for Plant Tissues, Munsell Color, Baltimore, Md., 1977. Light quality, photoperiod, and general growth of the plants affect color notations.
  • the internode stolon color of ⁇ Cavalier ⁇ stolons exposed to full sun is 5R 3/6.
  • Leaf blades of ⁇ Cavalier ⁇ are rolled in the bud, and are flat and stiff.
  • the leaf blade length of ⁇ Cavalier ⁇ ranges from 58.9 to 60.7 mm and from 1.33 to 1.79 mm in width (Table 3,4,5), shorter and narrower than ⁇ El Toro ⁇ and ⁇ Meyer ⁇ .
  • the genetic, adaxial leaf color of ⁇ Cavalier ⁇ is 2.5 GY 5/2 with ⁇ El Toro ⁇ having a leaf color of 2.5 GY 5/2, and ⁇ Meyer ⁇ having a color of 2.5 G 3/4.
  • the ligule of ⁇ Cavalier ⁇ is a row of silky hairs, approximately 2.1mm in length on the longest hairs.
  • the flat leaf length of ⁇ Cavalier ⁇ is a mean of 2.66 cm when measured in a greenhouse, Dallas, Tex., January 1996.
  • ⁇ Cavalier ⁇ has 2.5 GY 7/4 anthers and white colored stigmas, undistinguished in shade of color.
  • the inflorescence of ⁇ Cavalier ⁇ is a terminal spike-like raceme, with spikelets on short pedicels.
  • ⁇ Cavalier ⁇ has a mean culm length of 4.5 cm, a floral region of 13.3 mm, with a mean of 16.3 florets per raceme.
  • the chromosome number of ⁇ Cavalier ⁇ is 40.
  • ⁇ Cavalier ⁇ rated number one in the National Turf Evaluation Program (NTEP) trials over 4-yrs as tested at 24 locations ranging from California-Georgia and north to California and Colorado. ⁇ Cavalier ⁇ will find its primary utility in home lawns, golf course fairways and tee boxes and in parks and recreational areas (Table 6).
  • NEP National Turf Evaluation Program
  • ⁇ Cavalier ⁇ When ⁇ Cavalier ⁇ was compared with 59 other zoysiagrasses for salinity tolerance, it ranked number 22 in performance that was not different in injury rating for ⁇ Emerald ⁇ and superior to ⁇ Belair ⁇ , and ⁇ Meyer ⁇ (Table 8). When compared to other commercial varieties for root growth, ⁇ Cavalier ⁇ produced shorter average root depth, less root weight than ⁇ Emerald ⁇ , and clipping weights similar to ⁇ Emerald ⁇ (Table 9).
  • ⁇ Cavalier ⁇ is distinguished from other zoysiagrass by its fine texture, long-narrow leaf, with low rhizome, but high stolon production. It has good to excellent salt tolerance, and good shade tolerance. ⁇ Cavalier ⁇ is resistant to the fall armyworm and the tropical sod webworm. ⁇ Cavalier ⁇ is intermediate in its growth and recovery rate. ⁇ Cavalier ⁇ has good to excellent winter hardiness and will persist in regions north to Kansas, Missouri and Southern Illinois.
  • the zoysiagrass amplification profiles were obtained using primer of sequence GCCCGCCC, and are compared to the standard ⁇ Meyer ⁇ (FIG. 2). Complex banding patterns and amplification fragment length polymorphisms were obtained in all cases. Results indicate bands fall into two categories, those that are common to the species, and those that in combination are characteristic of the cultivar (some identified by dots).

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  • Health & Medical Sciences (AREA)
  • Life Sciences & Earth Sciences (AREA)
  • Natural Medicines & Medicinal Plants (AREA)
  • Physiology (AREA)
  • Botany (AREA)
  • Developmental Biology & Embryology (AREA)
  • Environmental Sciences (AREA)
  • Breeding Of Plants And Reproduction By Means Of Culturing (AREA)

Abstract

An asexually reproduced variety of perennial zoysiagrass with a unique combination of characters including an absence of leaf blade hairs, high turf quality, resistance to fall armyworm and a distinct DNA fingerprint.

Description

BRIEF SUMMARY OF THE INVENTION
The present invention relates to a new and distinct asexually reproduced variety of perennial zoysiagrass (Zoysia matrella (L.)) Merr.
BACKGROUND OF THE INVENTION
This invention relates to a new and distinct perennial zoysiagrass cultivar identified as `Cavalier` zoysiagrass (hereinafter referred to as `Cavalier`), that was tested as DALZ8507. `Cavalier` was discovered in a cultivated area that received mowing and traffic on Seoul National University campus, Suwon, South Korea. It was identified as a unique turfgrass clone growing among a diverse population of common zoysiagrasses native to the area.
For purposes of registration under the "International Convention for the Protection of New Varieties of Plants" (generally known by its French acronym as the UPOV Convention) and noting Section 1612 of the Manual of Plant Examining Procedure, it is proposed that the new variety of zoysiagrass of the present invention be named `Cavalier` Zoysiagrass.
BRIEF DESCRIPTIONS OF THE ILLUSTRATIONS
FIG. 1 depicts an up-close view of the leaf blade and ligule of `Cavalier`.
FIG. 2 depicts entire leaf blades and ligule of `Cavalier`.
FIG. 3 is a DNA fingerprint of `Cavalier` in comparison with known varieties of zoysiagrass.
DETAILED DESCRIPTION OF THE PLANT
`Cavalier` was characterized in greenhouse and field conditions. `Cavalier` is a unique variety of zoysiagrass (Zoysia matrella (L.) Merr that was discovered under cultivated conditions described above. `Cavalier` was asexually reproduced by cutting of stolons and rhizomes, rooting them in soil, and planting of the rooted material to provide planting stock for studying performance and for comparison of morphological characters after propagation. `Cavalier` has been propagated by sod, plugs, sprigs, and stolons. Seed reproduction with self-fertility is not common in the Zoysia sp. No seedling establishment from `Cavalier` has been noticed in either greenhouse or field studies.
`Cavalier` spreads primarily by stolons. It has an intermediate to rapid growth rate, and has intermediate to low water use requirements. `Cavalier` produces little thatch with an optimum mowing height of 3/8" to 2.5". `Cavalier` will generally cover in 10-12 months growing time when plug planted with 3"×4" plugs planted on 12" centers or by sprigs.
`Cavalier` has both rhizome and stolon growth. The stolons of `Cavalier` have a mean internode length of 16.7 mm between the fourth to the fifth node, 18.4 mm between the second and third nodes, with a mean stolon width and diameter from 1.35 to 1.14 mm (Tables 1,2). The stolons of `Cavalier` root adventitiously at the nodes. Color notations of plant tissues were based on the Munsell Color Charts for Plant Tissues, Munsell Color, Baltimore, Md., 1977. Light quality, photoperiod, and general growth of the plants affect color notations. The internode stolon color of `Cavalier` stolons exposed to full sun is 5R 3/6.
Leaf blades of `Cavalier` are rolled in the bud, and are flat and stiff. The leaf blade length of `Cavalier` ranges from 58.9 to 60.7 mm and from 1.33 to 1.79 mm in width (Table 3,4,5), shorter and narrower than `El Toro` and `Meyer`. There are not any hairs on the abaxial/adaxial leaf surfaces of `Cavalier`. Measured under greenhouse conditions in January 1996, the genetic, adaxial leaf color of `Cavalier` is 2.5 GY 5/2 with `El Toro` having a leaf color of 2.5 GY 5/2, and `Meyer` having a color of 2.5 G 3/4. The ligule of `Cavalier` is a row of silky hairs, approximately 2.1mm in length on the longest hairs.
The flat leaf length of `Cavalier` is a mean of 2.66 cm when measured in a greenhouse, Dallas, Tex., January 1996. `Cavalier` has 2.5 GY 7/4 anthers and white colored stigmas, undistinguished in shade of color. The inflorescence of `Cavalier` is a terminal spike-like raceme, with spikelets on short pedicels. `Cavalier` has a mean culm length of 4.5 cm, a floral region of 13.3 mm, with a mean of 16.3 florets per raceme.
The chromosome number of `Cavalier` is 40.
`Cavalier` rated number one in the National Turf Evaluation Program (NTEP) trials over 4-yrs as tested at 24 locations ranging from California-Georgia and north to Nebraska and Colorado. `Cavalier` will find its primary utility in home lawns, golf course fairways and tee boxes and in parks and recreational areas (Table 6).
This test was planted into a sited shaded with live oak trees in 1 Sep,. 1992. Under tree shade with competition from trees for available moisture and nutrients, `Cavalier` had slightly more cover than `Belair` and `Meyer` (Table 7).
When `Cavalier` was compared with 59 other zoysiagrasses for salinity tolerance, it ranked number 22 in performance that was not different in injury rating for `Emerald` and superior to `Belair`, and `Meyer` (Table 8). When compared to other commercial varieties for root growth, `Cavalier` produced shorter average root depth, less root weight than `Emerald`, and clipping weights similar to `Emerald` (Table 9).
Nine zoysiagrass cultivars were tested for resistance to the fall armyworm (Spodoptera frugiperda) in laboratory tests (Table 10). Eggs of the fall armyworm (FAW) were obtained from the colony maintained at the USDA Laboratory at Tifton, Ga. Larvae were developed from the eggs, and feed with zoysiagrass tissue of the cultivars, with days to pupation and adult emergence of the FAW used as indicators of FAW resistance. When neonate larvae of FAW were confined on the zoysiagrass cultivars, less than 5% of the larvae survived beyond 4 days of `Cavalier`, `Emerald`, or `Belair`. Two cultivars, `Palisades` and `8516` were among the most susceptible with 57% and 55% survival respectively to 17 days and longer. No larvae were able to survive to 17 days on the cultivar `Cavalier`, indicating the resistance of `Cavalier` to FAW was in the form of larval mortality and an extended development period (Table 10).
`Cavalier` is distinguished from other zoysiagrass by its fine texture, long-narrow leaf, with low rhizome, but high stolon production. It has good to excellent salt tolerance, and good shade tolerance. `Cavalier` is resistant to the fall armyworm and the tropical sod webworm. `Cavalier` is intermediate in its growth and recovery rate. `Cavalier` has good to excellent winter hardiness and will persist in regions north to Kansas, Missouri and Southern Illinois.
              TABLE 1
______________________________________
Internode length as measured between the second and third nodes,
internode diameter of the third internode, and node diameter of the
third node measured on zoysiagrass plants. Plants were grown in a
growth chamber with a 14-hour daylength, March 1995.
           Internode   Internode  node
           length      diameter   diameter
Genotype
mm--
mm--
mm--
______________________________________
`Cavalier` 18.4a*      1.35abc    1.35abc
`El Toro`  39.8a       1.45abc    1.45abc
`Meyer`    24.1a       1.99a      1.99a
______________________________________
 *Analysis of variance by General Linear Models, with means followed by th
 same letter not significantly different using Tukey's Studentized Range
 (HSD), alpha = 0.05. Only selected means presented.

              TABLE 2
______________________________________
Zoysiagrass internode lengths and stolen width measurements from the
fourth to the fifth nodes taken February 1988 on greenhouse grown
plants.
               Internode    Stolon
               length       width
Genotype
mm--
mm--
______________________________________
`Cavalier`     16.73bc*     1.14efg
`Emerald`       6.21e       1.02ghi
`Meyer`        16.47bc      1.53b
______________________________________
 *Analysis of variance by General Linear Models, with means followed by th
 same letter not significantly different using Tukey's Studentized Range
 (HSD), alpha = 0.05. Only selected means presented.

              TABLE 3
______________________________________
Zoysiagrass leaf measurements taken February 1988 on the fourth
youngest leaf from greenhouse grown plants.
              Blade width   Blade length
Genotype
mm--
mm--
______________________________________
`Cavalier`     1.79ij*      58.87e
`Emerald`      2.01fgh      30.56ff
`Meyer`       33.3b         82.33d
______________________________________
 *Means followed by the same letter are not significantly different using
 the WallerDuncan k ratio test (k ratio = 100). Only selected means
 presented.

              TABLE 4
______________________________________
Leaf blade width and length measured on the third youngest leaf of
zoysiagrasses. Plants were growing in a growth chamber with a
14-hour daylength, March 1995.
              Blade width   Blade length
Genotype
mm--
mm--
______________________________________
`Cavalier`    1.33d*        60.7cd
`El Toro`     3.26abc       68.3bcd
`Meyer`       2.59c         74.3bc
______________________________________
 *Analysis of variance by General Linear Models, with means followed by th
 same letter not significantly different using Tukey's Studentized Range
 (HSD), alpha = 0.05. Only selected means presented.

              TABLE 5
______________________________________
Zoysiagrass leaf sheath length measurements taken February 1988 on
the fourth youngest leaf from greenhouse grown plants.
                    Sheath length
Genotype
mm--
______________________________________
`Cavalier`          28.90ef*
`Emerald`           16.31ij
`Meyer`             30.68ed
______________________________________
 *Means followed by the same letter are not significantly different using
 the WallerDuncan k ratio test (k ratio = 100). Only selected means
 presented.

              TABLE 6
______________________________________
Mean turfgrass quality ranges of vegetative zoysiagrass cultivars grown
in the National Turfgrass Evaluation Program at 23 locations in the US
for 1992, 1993, and 1994.
                                       Overall
Variety      1992    1993   1994  1995 4-Yr Avg.
______________________________________
`Cavalier` (DALZ8507)
             5.95    6.23   5.89  5.99 5.93
`Marquis` (TC 2033)
             5.85    6.10   6.11  5.96 5.91
`Sunburst`   5.83    5.91   5.81  5.87 5.85
`TC 5018`    5.80    5.81   5.92  5.70 5.81
`Emerald`    5.74    6.21   6.05  5.73 5.79
`Omni` (CD 2013)
             5.56    6.13   6.06  5.96 5.73
`QT 2004`    5.63    6.01   5.86  5.57 5.63
`DALZ 8508`  5.59    6.06   5.74  5.60 5.60
`Palisades` (DALZ8514)
             5.82    5.82   5.46  5.44 5.59
`Royal` (DALZ9006)
             5.65    6.05   5.59  5.54 5.59
`Crowne` (DALZ8512)
             5.80    5.76   5.50  5.45 5.55
`El Toro`    5.78    5.63   5.34  5.41 5.50
`CD 259-13`  5.30    5.53   5.74  5.49 5.40
`Meyer`      5.26    5.70   5.76  5.47 5.39
`QT 2047`    5.37    5.39   5.26  5.16 5.30
`Belair`     4.99    5.58   5.61  5.02 5.16
`DALZ 8516`  4.72    5.42   4.96  5.05 4.86
`Diamond` (DALZ8502)
             4.40    5.03   4.58  4.36 4.41
`DALZ 8501`  4.88    4.31   3.99  4.05 4.27
`DALZ 8701`  4.23    4.10   3.71  3.58 3.85
LSD VALUE    0.22    0.20   0.20  0.20 0.17
______________________________________
 To determine statistical differences among entries, subtract one entry's
 mean from another entry's mean. Statistical difference occurs when this
 value is larger than the corresponding LSD value (LSD 0.05).
 In National Zoysiagrass Test  1991. Final Report 1992-95, NTEP No. 9615;
 (Table 4); United States Department of Agriculture, Agricultural Research
 Service, Beltsville Agricultural Research Center, Beltsville, MD 20705.

              TABLE 7
______________________________________
Mean turf cover, as percentage of plot during turf, during winter
1993-1994 for the 1991 NTEP zoysia trial planted under 80% shade in
Dallas, TX.
       Percentage Turf Cover
Variety  10 Nov 93    21 Dec 93
                               22 March 94
______________________________________
`Belair` 40.0         43.3     21.7
`Cavalier`
         48.3         55.0     26.7
`Emerald`
         41.7         51.7     26.7
`El Toro`
         35.0         31.7     21.7
`Meyer`  33.3         38.3     23.3
MSD      ns           14.5     10.2
______________________________________
 MSD is the minimum significant difference between entry means for
 comparison within column, and was based on the Duncan Waller kratio test
 (kratio = 100). Only selected means presented.
 In Morton, S. J., M. C. Engelke, and K. G. Porter. 1994. Performance of
 three warmseason turfgrass genera cultured in shade III. Zoysia spp. In
 Texas Turfgrass Research Report  1994. PR. 5242. p 27-29.

              TABLE 8*
______________________________________
Average percent shoot salt injury (average of 20 rating dates) on
zoysiagrass entries in the 1991 NTEP.sup.† Trials.
Entry         Source    Species.sup.‡
                                  % Injury
______________________________________
`Diamond`     NTEP 20   matrella  33ab.sup.
`El Toro`     NTEP 13   japonica  38a-e
`Emerald`     NTEP 10   jap x tenu
                                  42a-h
`Cavalier`    NTEP 17   matrella  42a-h
`Belair`      NTEP 11   japonica  50f-k
`Meyer`       NTEP 09   japonica  58lmn
`Korean Common`
              NTEP 07*  japonica  76pq
______________________________________
 *Selected data set; complete data set includes 59 varieties and cultivars
 .sup.† NTEP = National Turfgrass Evaluation Program.
 .sup.‡ Species identity.
 .sup..English Pound. Means followed by the same letter are not
 significantly different, based on the WallerDuncan kratio ttest (kratio =
 100)
 In Marcum, K. B., M. C. Engelke, S. J. Morton and C. Dayton. 1994.
 Salinity tolerances of selected bermudagrass and zoysiagrass genotypes. T
 Turfgrass Res.  1993, Consolidated Prog. Rep. PR 5140: 105-107.

              TABLE 9
______________________________________
Average mean root depth of zoysiagrasses grown in flexible tubes in
greenhouse studies, Dallas, TX.
         Average Mean  Total Root
                                Clipping
         Root Depth    Weight   Weight
Variety
mm--
mg--
mg--
______________________________________
`Belair` 296           330      286
`Cavalier`
         255           278      243
`El Toro`
         356           473      391
`Emerald`
         330           461      241
`Meyer`  333           411      466
MSD       79           161      267
______________________________________
 *MSD = minimum significant difference for comparison of means within
 columns based on the WallerDuncan kratio test where k = 100. Only selecte
 means presented.
 In Marcum, K. B., M. C. Engelke, S. J. Morton, and R. H. White. 1995.
 Rooting charcteristics and associated drought resistance of zoysiagrasses
 Agron. J. 87: 534-538.

              TABLE 10
______________________________________
Resistance in zoysiagrass to 4-day old larvae of the fall armyworm;
survival, weight, and development time.
Cultivars
        7 day larva   Pupa        Adult
and     alive  wt      alive
                            wt          alive
Genotypes
        %.sup.1
               mg.sup.2
                       %.sup.3
                            mg.sup.4
                                  days.sup.5
                                        %.sup.6
                                             days.sup.7
______________________________________
`Cavalier`
         0     --       0   --    --    0    --
`DALZ8501`
        14.8   34.9a.sup.8
                       11.1 151.2ab
                                  30.0a 11.1 40.0a
`K.Common`
        14.8   41.2ab  11.1 136.1ab
                                  30.3a 11.1 40.7a
`Belair`
        25.9   75.5b   25.9 149.8ab
                                  27.7ab
                                        25.9 37.6abc
`El Toro`
        33.3   58.7ab  26.7 171.1a
                                  29.0a 26.7 39.0a
`Emerald`
        37.0   52.1ab  37.0 137.5ab
                                  30.5a 37.0 40.5a
`Meyer` 60.0   55.4ab  53.1 148.3ab
                                  28.9a 53.3 38.8ab
`Palisades`
        66.7   131.6c  66.7 122.7b
                                  25.2b 66.7 35.3bc
`DALZ8516`
        81.5   143.9c  74.1 156.7ab
                                  25.1b 74.1 34.7c
______________________________________
 .sup.1 Number of surviving 17day-old-larvae.
 .sup.2 Mean larval weight taken after 17 days of feeding.
 .sup.3 Number surviving to pupation.
 .sup.4 Mean weight of pupa within 2 days of pupation.
 .sup.5 Mean number of days to pupation.
 .sup.6 Mean survival to adult.
 .sup.7 Mean number of days from egg hatch to adult emergence.
 .sup.8 Means in a column followed by the same letter are not significantl
 different by WallerDuncan kratio t test (k = 100) (P = 0.05)
 In Reinert, J. A., M. C. Engelke, S. J. Morton, P. S. Graff, and B. R.
 Wiseman. 1994. Resistance in zoysiagrass (Zoysia spp) to the fall army
 worm (Spodoptera frugiperda). TX turfgrass Research1992. Consolidated
 Prog. Rep. PR 5248: 39-4
EXAMPLE 1 DNA Fingerprint Analysis
See Caetano-Annolles, B. J. Bassam and Peter M. Gresshoff, 1991, DNA Amplification fingerprinting using very short arbitrary oligonucleotide primers. Biotechnology. Vol. 9. Pp. 553-557.
The zoysiagrass amplification profiles were obtained using primer of sequence GCCCGCCC, and are compared to the standard `Meyer` (FIG. 2). Complex banding patterns and amplification fragment length polymorphisms were obtained in all cases. Results indicate bands fall into two categories, those that are common to the species, and those that in combination are characteristic of the cultivar (some identified by dots).
Statistical Analysis
Statistical analysis performed utilizes the method set forth in Steel and Torrie, 1960, Principles and Procedures of Statistics. Pages 1-481. McGraw-Hill Book Company, Inc. New York.

Claims (1)

I claim:
1. A new and distinct variety of zoysiagrass (Zoysia matrella (L.) Merr.) plant as described and illustrated herein, having the principle distinguishing characteristics of white stigmas, and absence of leaf blade hairs, high turf quality, resistance to fall armyworm and a distinct DNA fingerprint.
US08/890,959 1997-07-08 1997-07-08 `Cavalier` zoysiagrass plant Expired - Lifetime USPP10778P (en)

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US08/890,959 USPP10778P (en) 1997-07-08 1997-07-08 `Cavalier` zoysiagrass plant
PCT/US1998/014283 WO1999002029A1 (en) 1997-07-08 1998-07-08 Cavalier zoysiagrass plant
JP2000501646A JP2001509363A (en) 1997-07-08 1998-07-08 Caverns plant

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US08/890,959 USPP10778P (en) 1997-07-08 1997-07-08 `Cavalier` zoysiagrass plant

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Cited By (6)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
USPP11466P (en) * 1998-02-26 2000-08-01 Sod Solutions, Inc. Zoysia grass plant named `SS-500`
USPP11495P (en) * 1998-02-24 2000-08-29 Sod Solutions, Inc. Zoysia grass plant named `SS-300`
USPP22014P3 (en) 2009-05-14 2011-07-05 GeneGro Pty Ltd Zoysiagrass plant ‘A-1’
USPP25203P3 (en) * 2013-04-16 2014-12-30 David Doguet Zoysiagrass plant named ‘L1F’
USPP31464P3 (en) 2018-05-07 2020-02-18 The Texas A&M University System Zoysiagrass named ‘KSUZ 0802’
USPP35643P2 (en) 2023-02-01 2024-02-06 David Doguet Zoysiagrass plant named ‘L1FS’

Citations (2)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
USPP5845P (en) * 1984-11-21 1986-12-30 The Regents Of The University Of California Zoysiagrass plant `El Toro`
USPP6529P (en) * 1987-03-18 1989-01-10 Zoysia grass plant cv `P-1`

Family Cites Families (2)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
USPP6516P (en) * 1987-03-16 1989-01-03 Zoysia grass plant (ZT-4)
USPP7074P (en) * 1988-12-09 1989-12-12 Zoysia Japonica grass plant ZT-11

Patent Citations (2)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
USPP5845P (en) * 1984-11-21 1986-12-30 The Regents Of The University Of California Zoysiagrass plant `El Toro`
USPP6529P (en) * 1987-03-18 1989-01-10 Zoysia grass plant cv `P-1`

Cited By (6)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
USPP11495P (en) * 1998-02-24 2000-08-29 Sod Solutions, Inc. Zoysia grass plant named `SS-300`
USPP11466P (en) * 1998-02-26 2000-08-01 Sod Solutions, Inc. Zoysia grass plant named `SS-500`
USPP22014P3 (en) 2009-05-14 2011-07-05 GeneGro Pty Ltd Zoysiagrass plant ‘A-1’
USPP25203P3 (en) * 2013-04-16 2014-12-30 David Doguet Zoysiagrass plant named ‘L1F’
USPP31464P3 (en) 2018-05-07 2020-02-18 The Texas A&M University System Zoysiagrass named ‘KSUZ 0802’
USPP35643P2 (en) 2023-02-01 2024-02-06 David Doguet Zoysiagrass plant named ‘L1FS’

Also Published As

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JP2001509363A (en) 2001-07-24
WO1999002029A1 (en) 1999-01-21

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